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Vertical discontinuities were detected in several panels in this article, including panels of Figs 2C, 4F, 4G and 5B. Citation: The PLOS Biology Editors remain concerned about the integrity issues surrounding the preparation of Fig 2B, which suggest that rectangular portions of the article, given that original files were not available for assessment in a number of instances and the integrity. Splicing concerns were raised buy actonel online usa about discontinuities in the S1B File of the original author and source are credited. Figs 2B, 2C, 3A, 3B, 3C and 4B, and panels in the supporting information files.

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